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1、GeneGenealogies,VariationandEvolutionThispageintentionallyleftblankGeneGenealogies,VariationandEvolutionAPrimerinCoalescentTheoryJotunHeinUniversityofOxford,UKMikkelH.SchierupandCarstenWiufUniversityofAarhus,Denmark13GreatClarendonStreet,OxfordOX26DPOxford
2、UniversityPressisadepartmentoftheUniversityofOxford.ItfurtherstheUniversity’sobjectiveofexcellenceinresearch,scholarship,andeducationbypublishingworldwideinOxfordNewYorkAucklandBangkokBuenosAiresCapeTownChennaiDaresSalaamDelhiHongKongIstanbulKarachiKolkata
3、KualaLumpurMadridMelbourneMexicoCityMumbaiNairobiSãoPauloShanghaiTaipeiTokyoTorontoOxfordisaregisteredtrademarkofOxfordUniversityPressintheUKandincertainothercountriesPublishedintheUnitedStatesbyOxfordUniversityPressInc.,NewYork©JotunHein,MikkelH.Schierupa
4、ndCarstenWiuf,2005ThemoralrightsoftheauthorshavebeenassertedDatabaserightOxfordUniversityPress(maker)Firstpublished2005Allrightsreserved.Nopartofthispublicationmaybereproduced,storedinaretrievalsystem,ortransmitted,inanyformorbyanymeans,withoutthepriorperm
5、issioninwritingofOxfordUniversityPress,orasexpresslypermittedbylaw,orundertermsagreedwiththeappropriatereprographicsrightsorganization.EnquiriesconcerningreproductionoutsidethescopeoftheaboveshouldbesenttotheRightsDepartment,OxfordUniversityPress,attheaddr
6、essaboveYoumustnotcirculatethisbookinanyotherbindingorcoverandyoumustimposethissameconditiononanyacquirerBritishLibraryCataloguinginPublicationDataDataavailableLibraryofCongressCataloginginPublicationDataDataavailableISBN0-19-852995-3(hbk.)ISBN0-19-852996-
7、1(pbk.)13579108642TypesetbyNewgenImagingSystems(P)Ltd.,Chennai,IndiaPrintedinGreatBritainonacid-freepaperbyBiddlesLtd.,King’sLynn,NorfolkPrefaceCoalescenttheoryhasgonefromanobscurecornerofpopulationgeneticstoacentralconceptforanybodythatstudiesvariationatt
8、hesequencelevel.Besidesfillingtheobviousneedforabookonthissubject,itisalsoourwishtopresentthistheoryinastraightforwardandelementarymannerthatcoulddispelthemisconceptionthatcoalescenttheoryisinherentlyverydiffic