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ID:44156699
大小:4.28 MB
页数:146页
时间:2019-10-19
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1、cyclicLesson9Theoriginofspecies1.ThebiologistsdefineaspeciesModernbiologistsgenerallydefineaspeciesasgroupofactuallyorpotentiallyinterbreedingpopulationsthatisreproductivelyisolatedfromothersuchgroups.Membersofspeciescaninterbreedwitheachother,buttheycannotbreedwithorganismsbelongingtoanothe
2、rspecies.Oneadvantageofthestandardofreproductiveisolationisthatitisveryprecise.Notice,however,thatitcanonlybeappliedtoorganismsthatreproducesexually.Asexualreproducers,includingmostprokaryotes,manyplants,andsomeanimals,mustbeclassifiedintospeciesonthebasisofphysical(biochemicalormorphologica
3、l)traits.2.PreventinggeneexchangeTwogeneraltypesofmechanismsoperatetoblocktheexchangeofgenesbetweenindividualsofrelatedgroups.Thefirstgeneraltypeismadeupofprezygoticisolatingmechanisms-mechanismsthatpreventtheformationofzygotes.Prezygoticisolationfallsintotwocategories:ecologicalandbehaviora
4、l.Inthefirstcase,tworelatedgroupmaybecomeadaptedtoslightlydifferentenvironments_perhapsvaryingsoiltypesorfoodsources.Overtime,thesegeneticdifferencesbecomesogreatthatsuccessfulcross-fertilizationcannolongertakeplace.Inbehavioralisolation,relatedgroupsevolvedifferingbehaviors-suchasspeciesmat
5、ingrituals-thatrestricttheexchangeofgenestomembersofthesamegroup.-----PreventinggeneexchangeSometimesthedifferencesthatproduceprezygoticisolationinvolvemechanicalisolation.Thatis,matingisphysicallyimpossiblebetweenmembersofdifferentspeciesbecausegenitalsofmalesandfemalesarestructurallyincomp
6、atibleorbecausemoleculesonthesurfacesofspermandeggfailtobind.Afinaltypeofprezygoticmechanismistemporalisolation,inwhichtime-relatedenvironmentalcuesthattriggerreproductiveprocessesaredifferentforrelatedspecies.Inpostzygoticisolatingmechanismsmatingoccurs,buttheresultinghybridorganismisinviab
7、leorsterile.Inaspecialcaseofhybridsterilitytermedhybridbreakdown,thesecondandsubsequentgenerationsafteracrossshowreducedreproductivesuccess.Inpostzygoticisolatingmechanismsmatingoccurs,buttheresultinghybridorganismisinviableorsterile.Inaspecialcase
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