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时间:2020-03-15
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1、Ithaslongbeenknownthattherateofoxidativemetabolism(theprocessthatusesoxygentoconvertfoodintoenergy)inanyanimalhasaprofoundeffectonitslivingpatterns.Thehighmetabolicrateofsmallanimals,forexample,givesthemsustainedpowerandactivityperunitofweight,butatthecostofrequiringconstantconsumptionoff
2、oodandwater.Verylargeanimals,withtheirrelativelylowmetabolicrates,cansurvivewellonasporadicfoodsupply,butcangen-eratelittlemetabolicenergypergramofbodyweight.Ifonlyoxidativemetabolicrateisconsidered,there-fore,onemightassumethatsmaller,moreactive,animalscouldpreyonlargerones,atleastifthey
3、attackedingroups.Perhapstheycouldifitwerenotforanaerobicglycolysis,thegreatequalizer. Anaerobicglcolysisisaprocessinwhichenergyisproduced,withoutoxygen,throughthebreakdownofmuscleglycogenintolacticacidandadenosinetri-phosphate(ATP),theenergyprovider.Theamountofenergythatcanbeproducedanae
4、robicallyisafunctionoftheamountofglycogenpresent-inallvertebratesabout0.5percentoftheirmuscles'wetweight.Thustheanaerobicenergyreservesofaverte-brateareproportionaltothesizeoftheanimal.If,forexample,somepredatorshadattackeda100-tondinosaur,normallytorpid,thedinosaurwouldhavebeenabletogene
5、ratealmostinstantaneously,viaanaerobicglycolysis,theenergyof3,000humansatmaximumoxidativemetabolicenergyproduction.Thisexplainshowmanylargespecieshavemanagedtocompetewiththeirmoreactiveneighbors:thecompensationforalowoxidativemetabolicrateisglycolysis. Therearelimitations,however,tothisc
6、ompensa-tion.Theglycogenreservesofanyanimalaregood,atmost,foronlyabouttwominutesatmaximumeffort,afterwhichonlythenormaloxidativemetabolicsourceofenergyremains.Withtheconclusionofaburstofactivity,thelacticacidlevelishighintthebodyfluids,leavingthelargeanimalvulnerabletoattackuntiltheacidis
7、reconverted,viaoxidativemetabolism,bytheliverintoglucose,whichisthensent(inpart)backtothemusclesforglycogenresyn-thesis.Duringthisprocesstheenormousenergydebtthattheanimalhasrunupthroughanaerobicglycolysismustberepaid,adebtthatisproportionallymuchgreaterforthelarger
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