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1、Eur.J.Entomol.104:303–309,2007http://www.eje.cz/scripts/viewabstract.php?abstract=1232ISSN1210-5759CleorodesWarren,1894doesnotbelonginthetribeBoarmiini(Lepidoptera:Geometridae)12343,5JAANVIIDALEPP,TOOMASTAMMARU*,NIINASNÄLL,KAIRUOHOMÄKIandNIKLASWAHLBERG1InstituteofAgriculturalandEnvi
2、ronmentalSciences,EstonianUniversityofLifeSciences,Kreutzwaldi64,EE-51014Tartu,Estonia2InstituteofZoologyandHydrobiology,UniversityofTartu,Vanemuise46,EE-51014Tartu,Estonia3LaboratoryofGenetics,DepartmentofBiology,UniversityofTurku,FI-20014Turku,Finland4SectionofEcology,Departmentof
3、Biology,UniversityofTurku,FI-20014Turku,Finland5DepartmentofZoology,StockholmUniversity,S-10691Stockholm,SwedenKeywords.Taxonomy,morphology,molecularphylogeny,directoptimisation,Geometridae,Ennominae,Gnophini,Boarmiini,Bistonini,CleorodeslichenariaAbstract.ThegeometridgenusCleorodes
4、isshowntobelonginthetribeGnophini(sensulato)andnotinBoarmiiniaspreviouslyassumed.Theconclusionisbasedonananalysisofmorphologicalcharactersofanumberofgenerainthesetribes.Moreover,theresultisunambiguouslysupportedbyaphylogeneticanalysisofDNAsequencevariationinthreenucleargeneregions(s
5、egmentsD1andD2of28SrRNA,andelongationfactor1a)andamitochondrialgene,cytochromeoxidase-1.Thephylogenetichypothesisisbasedonacombinedsequencedataset,whichwasanalysedusingdirectoptimisation.INTRODUCTIONEnnominae.Thesemothsarerelativelyuniforminappearance,withthewingpatternresemblingtre
6、ebark,ThelargestsubfamilyofGeometridae,theEnnominae,andhavetraditionallybeenviewedasanuncontroversialwithover9700describedspecies(about45%ofallGeo-naturalunit.Somedifferentlylookinggenerahavebeenmetridae:Minet&Scoble,1999),iswelldefinedbyaaddedtothetribe(e.g.Arichanna,byHerbulot,196
7、2),morphologicalcharacter,theweakeningorabsenceofthebutthesystematicpositionofanyofthe“classicalBoar-veinM2inthehindwings.Ontheotherhand,thereisnomias”hasneverbeenquestioned,neitherhasthetribeasaconsensusaboutthetaxonomicaffinitiesofthenumerouswholebeensubjectedtoasystematictaxonomi
8、crevision.currently