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1、中国水稻科学(ChinJRiceSci),２０１６,３０(１):９９－１０５http://www．ricesci．cnDOI:１０．１６８１９/j．１００１Ｇ７２１６．２０１６．５１５３９９水稻小穗颖壳发育的研究进展徐乾坤＃任德勇＃李自壮曾大力郭龙彪钱前∗(中国水稻研究所水稻生物学国家重点实验室,杭州３１０００６;＃共同第一作者;∗通讯联系人,EＧmail:qianqian１８８＠hotmail．com)ResearchProgressesinRiceSpikeletGlumeDevelopmentXUQianＧkun＃,RENDeＧyong＃,LIZiＧzhuang,ZENGDa
2、Ｇli,GUOLongＧbiao,QIANQian∗(StateKeyLaboratoryofRiceBiology,ChinaNationalRiceResearchInstitute,Hangzhou３１０００６,China;＃Theseauthorscontributedequallytothiswork;∗Correspondingauthor,EＧmail:qianqian１８８＠hotmailc．om)XUQiankun,RENDeyong,LIZizhuang,etal．Researchprogressesinricespikeletglumedevelopment．
3、ChinJRiceSci,２０１６,３０(１):９９Ｇ１０５．Abstract:Rice(OryzasativaL．),amonocotmodelplant,isanimportantcerealcropintheworld．Thefloweringtime,inflorescenceandfloralorganmorphologicalstructurehavesignificantinfluenceonriceyieldandquality．ThereＧsearchonthestructureanddevelopmentoffloralorgansishelpfultoimpr
4、ovethegrainyieldandricequality．ThedeＧvelopmentandmorphogenesisoffloralorganisavitalprocessfromvegetativegrowthtoreproductivegrowthinrice．Moreandmorebiologicalresearchesfocusonitsdevelopmentalpatternandmolecularmechanism．Theglumesofricespikeletsareuniqueorgansandconsistedoflemmas,paleae,sterile
5、lemmasandrudimentaryglumes．Themolecularmechanismsoftheformationandoriginofglumeskeeppoorunderstanding．Recently,furtherstudyoftheglumeshelptonotonlyunderstandthericespikeletandfloralorgandevelopment,butalsofacilitateunderstandingoftheregulatorynetworkinvolvedinricespikeletandfloralorgandevelopm
6、ent．Inthispaper,wefocusonthericeglumedevelopmentandreviewtheABCDEmodeloffloralorganspecialization．Keywords:rice(OryzasativaL．);paleaandlemma;sterilelemma;rudimentaryglume;ABCDEmodel徐乾坤,任德勇,李自壮,等．水稻小穗颖壳发育的研究进展．中国水稻科学,２０１６,３０(１):９９Ｇ１０５．摘要:水稻(OryzasativaL．)是世界上重要的粮食作物,也是单子叶植物的模式植物.开花时间、花序和花器官的形态结
7、构对其产量和品质均有重要影响.对花器官形态结构及发育机理的研究有助于提高水稻产量并改良其品质.花器官的形成和发育是水稻从营养生长转向生殖生长的重要过程,其发育模式和分子机理,一直是生物学研究的热点和焦点.水稻小穗的颖壳是禾本科特有的器官,主要包括内外稃、护颖和副护颖,关于其起源和形成的分子机制还知之甚少.近些年对颖壳的研究不断深入,不仅有助于深入认识水稻小穗或花器官的发育,而且能系统地了解水稻小穗或花器官发育的整个调控网络.本文主要介绍了水稻小穗颖壳发育的相关进展及植
