高级植物生理学 课件 第七章 植物的生殖生理

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1、1PlantReproductionZichaoMao2Lifecycleofplant3TransitiontoreproductionFlowerorgandevelopmentGametogenesisandfertilizationPlantReproduction4TransitiontoreproductionVegetativephaseReproductivephase?InflorescenceFlower5ProductionofflowersinvolvestwotransitionsinArabidopsisSC:st

2、emcellP:organprimordiaSe:sepalConvertSAM（ShootApicalMeristem)toinflorescencemeristem(infinite,makinglateralorgans)2.Convertinflorescencemeristemtofloralmeristem(terminal,flowers)6FactorsregulatingthetransitionsVegetativemeristemInflorescencemeristemFloralmeristemGenes(flowe

3、ring-timegenesandfloralidentitygenes)Light(photoperiod)ThebiologicalclockTemperatureHormones7affectingthetransitionofvegetativegrowthtoreproductivegrowthWTemf2Flowering-timegenesemf1embryonicflower8affectingformationofinflorescenceandfloralmeristemsFloralidentitygenesInflor

4、escence(fromInflorescencemeristem)Flower(fromFloralmeristem)9terminalflower1(tfl1):Converttheinflorescencemeristemtotheflowermeristem.leafy(lfy):producemoreinflorescences,delayedfloweringMutationsinfloralidentitygenestfl1[FUNCTION]Controlsinflorescencemeristemidentityandisr

5、equiredformaintenanceofaindeterminateinflorescence.Preventstheexpressionof'APETALA1'and'LEAFY'.Alsoplaysaroleintheregulationofthetimeoffloweringinthelong-dayfloweringpathway.Mayformcomplexeswithphosphorylatedligandsbyinterferingwithkinasesandtheireffectors]Expressedbelowthe

6、apicaldomeofinflorescenceandcoflorescencemeristems,andininflorescencestem.Weaklyexpressedinvegetativephasefromday2orday3.Increasedexpressionaftercommitmenttofloweringfromday7on.LEAFY;LFY,FUNCTIONSIN:chromatinDNAbinding,transcriptionfactoractivity,sequence-specificDNAbinding

7、;INVOLVEDIN:flowerdevelopment,maintenanceofinflorescencemeristemidentity,responsetogibberellinstimulus,gibberellicacidmediatedsignaling;LOCATEDIN:nucleus;EXPRESSEDIN:shootapex,leafwhorl,embryo,flower,seed;EXPRESSEDDURING:7growthstages;12Factorsregulatingthetransitiontorepro

8、ductionVegetativemeristemInflorescencemeristemFloralmeristemEMFTF1LFY13GarnerandAl