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1、ROUTLEDGELIBRARYEDITIONS:20THCENTURYSCIENCEVolume20THESTRATEGYOFTHEGENESThispageintentionallyleftblankTHESTRATEGYOFTHEGENESADiscussionofSomeAspectsofTheoreticalBiologyC.H.WADDINGTONFirstpublishedin1957Thiseditionfirstpublishedin2014byRoutledge2ParkSquare,Milton
2、Park,Abingdon,Oxon,OX144RNandbyRoutledge711ThirdAvenue,NewYork,NY10017RoutledgeisanimprintoftheTaylor&FrancisGroup,aninformabusiness©1957GeorgeAllen&UnwinLtd.Allrightsreserved.Nopartofthisbookmaybereprintedorreproducedorutilisedinanyformorbyanyelectronic,mecha
3、nical,orothermeans,nowknownorhereafterinvented,includingphotocopyingandrecording,orinanyinformationstorageorretrievalsystem,withoutpermissioninwritingfromthepublishers.Trademarknotice:Productorcorporatenamesmaybetrademarksorregisteredtrademarks,andareusedonlyf
4、oridentificationandexplanationwithoutintenttoinfringe.BritishLibraryCataloguinginPublicationDataAcataloguerecordforthisbookisavailablefromtheBritishLibraryISBN:978-0-415-73519-3(Set)eISBN:978-1-315-77941-6(Set)ISBN:978-1-138-01731-3(Volume20)eISBN:978-1-315-779
5、32-4(Volume20)Publisher’sNoteThepublisherhasgonetogreatlengthstoensurethequalityofthisbookbutpointsoutthatsomeimperfectionsfromtheoriginalmaybeapparent.DisclaimerThepublisherhasmadeeveryefforttotracecopyrightholdersandwouldwelcomecorrespondencefromthosetheyhave
6、beenunabletotrace.THESTRATEGYOFTHEGENESADiscussi011ofSomeAspectsofTheoreticalBiologyBYc.H.WADDINGTONSC.D.,F.R.S.WITHANAPPENDIXBYH.KAeSERPH.D.RuskinHouseGEORGEALLEN&UNWINLTDMUSEUMSTREETLONDONfIRSTPUBLISHEDIN1957ThisbookiscopyrightundertheBerneConvention.Apartfr
7、omanyfairdealingforthepurposesofprivatestudy,research,criticismorreview,aspermittedundertheCopyrightAct1911,noportionmaybereproducedbyanyprocesswithol4twrittenpermission.Enquiryshouldbemadetothepuhlisher©GeorgeAllen&UnwinLtd.,1957PRINTEDINGREATBRITAINin12point
8、BembotypeBYJ.W.ARROWSMITHLTD.WINTBRSTOKEROAD,BRISTOL3PREFACETHEFIVEessayswhichformthebodyofthisbook,andthesixthonewhichmyfriendDr.HenrikKacserhaskindlyallowedmetoinclude,areconcern