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1、EVOLUTIONANDTHELEVELSOFSELECTIONThispageintentionallyleftblankEvolutionandtheLevelsofSelectionSAMIROKASHACLARENDONPRESS·OXFORD1GreatClarendonStreet,Oxfordox26dpOxfordUniversityPressisadepartmentoftheUniversityofOxford.ItfurtherstheUniversity’sobjectiveofexcellenceinresearch,scholars
2、hip,andeducationbypublishingworldwideinOxfordNewYorkAucklandCapeTownDaresSalaamHongKongKarachiKualaLumpurMadridMelbourneMexicoCityNairobiNewDelhiShanghaiTaipeiTorontoWithofficesinArgentinaAustriaBrazilChileCzechRepublicFranceGreeceGuatemalaHungaryItalyJapanPolandPortugalSingaporeSout
3、hKoreaSwitzerlandThailandTurkeyUkraineVietnamOxfordisaregisteredtrademarkofOxfordUniversityPressintheUKandincertainothercountriesPublishedintheUnitedStatesbyOxfordUniversityPressInc.,NewYork©SamirOkasha2006ThemoralrightsoftheauthorhavebeenassertedDatabaserightOxfordUniversityPress(m
4、aker)Firstpublished2006Allrightsreserved.Nopartofthispublicationmaybereproduced,storedinaretrievalsystem,ortransmitted,inanyformorbyanymeans,withoutthepriorpermissioninwritingofOxfordUniversityPress,orasexpresslypermittedbylaw,orundertermsagreedwiththeappropriatereprographicsrightso
5、rganization.EnquiriesconcerningreproductionoutsidethescopeoftheaboveshouldbesenttotheRightsDepartment,OxfordUniversityPress,attheaddressaboveYoumustnotcirculatethisbookinanyotherbindingorcoverandyoumustimposethesameconditiononanyacquirerBritishLibraryCataloguinginPublicationDataData
6、availableLibraryofCongressCataloginginPublicationDataDataavailableTypesetbyLaserwordsPrivateLimited,Chennai,IndiaPrintedinGreatBritainonacid-freepaperbyBiddlesLtd,King’sLynn,NorfolkISBN0–19–926797–9978–0–19–926797–213579108642AcknowledgementsIamgratefultonumerousfriendsandcolleagues
7、fordiscussionandcorrespondence,inparticularPeterGodfrey-Smith,ElliottSober,DavidSloanWilson,JohnDamuth,KimSterelny,thelateJohnMaynardSmith,JimGriesemer,thelateBillHamilton,AyeletShavit,LisaLloyd,BenKerr,PatrickForber,KenReisman,EvaJablonka,DaveChalmers,RickMichod,RobWilson,TomHenfre
8、y,RasmusWinther,Bre
