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1、ONLINESUPPLEMENTARYMATERIAL1Contents1.FaunalList2.Stratigraphy3.SpeciesDescriptions4.PhylogeneticAnalysisa.RevisionstoGauthieretal.2012matrixb.NewCharacters/CharacterIllustrationsc.MaterialsStudiedd.Methodse.Resultsf.Discussion5.Resampling6.MorphometricAnalysis7.Phylogeneti

2、cIndependentContrastsAnalyses8.References9.Character-TaxonMatrix1.FaunalList,UpperMaastrichtianofWesternNorthAmericaBasedonliterature(1-6)andmuseumspecimens(seeSI3forcompletereferencesandspecimennumbers).Speciesinboldareeithernamedorrecognizedhereforthefirsttime.TableS1.IGU

3、ANIAIguanidaeincertaesedisPariguanalancensisPOLYGLYPHANODONTIAPolyglyphanodontiaincertaesedisObamadongracilisPolyglyphanodontidaePolyglyphanodonsternbergiChamopsiidaeChamopssegnisLeptochamopsdenticulatusMeniscognathusaltmaniHaptosphenusplacodonStypodontosaurusmelletesTripen

4、naculusn.sp.FrenchmanchamopsiidPeneteiusaquiloniusSocognathusbrachyodonLaramiechamopsiidSCINCOMORPHAScincomorphaincertaesedisLonchisaurustrichurusScincoideaincertaesedisEstescincosauruscookiGlobauridaeContogenyssloaniANGUIMORPHAXenosauridaeExostinuslancensisAnguidaeOdaxosau

5、ruspiger“Gerrhonotus”sp.PlatynotaincertaesedisLitakisgilmoreiColpodontosauruscracensPalaeosaniwasp.ParadermabogertiCemeteriusmonstrosusParasaniwawyomingensisOPHIDIAConiophidaeConiophisprecedensAlethinophidiaincertaesedisCerberophisrobustusLanceSnakeSQUAMATAINCERTAESEDISLami

6、asaurusferoxSweetwaterCountylizard2.StratigraphyA.MaastrichtianTheCretaceouslizardsincludedinthisstudycomefromtheHellCreekFormationofMontana,theLanceandFerrisformationsofWyoming,theFrenchmanFormationofSaskatchewan,theScollardFormationofAlberta,theLaramieFormationofColorado,

7、andtheNorthHornFormationofUtah.AllformationsareinferredtobelateMaastrichtianinageandcontaintypicallateMaastrichtianfaunas(i.e.,LancianLandVertebrateAge).Ageconstraint(TableS2)isbasedonseverallinesofevidence,includingradiometricdating,magnetostratigraphy,palynostratigraphy,a

8、ndvertebratebiostratigraphy,whichisprimarilybasedonmammalianfaunas.i.FrenchmanForm