4、 EugeniusWarming)的影响,明确提出生态系统的概念。认为:“(原文)Butthefundmentalconceptionis,asitseemstome,thewhole system (inthesenseofphysics),includingnotonlytheorganism-complex,butalsothewhilecomplexofphysicalfactorsformingwhatwecalltheenviriment,withwhichtheyformonephysicalsystem....These ecosystems,aswemayca